The Cactaceae family includes about 1,650 to 3,500 species of cacti and succulents classified in 130 genera. Because they live in harsh, arid environments, these fleshy, spiny perennial plants have developed a variety of unique characteristics for protection and to retain water, reduce evaporation, and resist heat.
The word “succulent” is derived from the Latin term sucus, meaning sap. All cacti are succulents. The word “cactus” is derived from the Greek term kaktos, describing thistles. Botanists estimate cacti first existed during the Mesozoic era, about 130 million years ago. Limited cacti fossil evidence exists (the earliest known specimen is about forty thousand years old).
Cacti vary in size. The Copiapoa laui is a spherical plant several millimeters in diameter, while the Pachycereus weberi is cylindrical, stands more than 20 meters tall, and can weigh more than 25 tons. Cacti often develop bizarre shapes to cope with arid conditions. Some stems are flat, and others are puffy. Many consist of jointed segments, while others have one round stem.
Cacti stems swell when storing water. Surface ridges and grooves gather water. Roots extend in a wide area near the soil surface, to capture any moisture. The pincushion, barrel, saguaro, prickly pear, night-blooming cereus, and Christmas cactus are some of the most familiar cacti.
Unlike other plants, cacti have areolas on stems where branches, spines, glochids (bristles), leaves, and flowers grow. Spines protect and shade the plant and its seedlings from predators and ultraviolet radiation and serve as condensation sites. Known as crassulacean acid metabolism (CAM), photosynthesis in cacti is reversed from the process in other plants. Stems have chlorophyll because leaves are either absent or tiny.
At night, instead of day, cacti open the stomata on their stems to collect carbon dioxide and expel oxygen. The carbon dioxide is stored as organic acids for conversion to sugar during the day. Because the temperature is cooler when the stomata are opened, less water is lost. During the day, the closed stomata prevent evaporation from occurring.
Life Cycles of Cacti
seeds if they are shaded and not consumed by predators.
Other cacti emerge from stems and take root where they fall. Artificially, cacti can also take root from cuttings. Diverse insects and animals are attracted to the flowers and assist in their pollination. Some birds nest in holes in cacti stems.
Distribution of Cacti
Cacti grow in deserts, prairies, mountains, and tropical climates and have developed a tolerance for extreme conditions. Cacti are indigenous to North, Central, and South America. The Epiphyllum species live in tropical trees. Other cacti grow in rocky places.
Some Chilean cacti in the Atacama Desert secure water from sea fog. The largest and most diverse population of cacti is in Mexico. The prickly pear is the most widely distributed cactus, ranging from near the Arctic circle to southern South America.
Uses of Cacti
medicinal purposes. While no cacti are poisonous, some species have unpleasant chemicals that discourage predators.
Some hybrids have naturally occurred, and cactus segments can be detached at joints to graft to artificially unique plants. Because of poaching, cacti are considered endangered plants, with some species threatened by extinction, and are federally protected at Saguaro National Park (in Arizona’s Tucson Basin) and Organ Pipe Cactus National Monument (in Arizona’s Sonoran Desert).
Yucca and jade plants are two of the most familiar succulents.
Because of evolutionary adaptation to endure climatic extremes, succulents have small leaves and spongy tissues that keep water for prolonged durations. Succulents retain water to with stand such environmental stresses as drought, scorching wind, shallow or salty topsoil, steep locations, and over-crowding by other plants.
Succulents keep flower stalks and fruit until all the water is depleted from them. They have a thick skin, which is waxy, and sometimes alter their shape while adjusting to differences of light and moisture. Most succulents are gray, although a few are colored lilac, pink, light green, beige, or ivory, often in patterns that may serve as camouflage.
The greatest quantity and most diverse succulents can be found in Mexico and South Africa, which have thousands of species. New species are still being discovered because of variations arising from environmentally triggered adaptations. Some succulents, such as the Argyroderma, are abundant, growing in thick clumps.
The rarer succulents include Conophytum burgeri, which lives on only one South African hill. Succulents are threatened by overgrazing and industrial and agricultural development of habitats. Some succulents, particularly aloe, have healing juices to soothe burns.